Logistic Models

Exponential growth can't last. A population that outgrows its food or space slows down, and its size levels off at the most the environment can support. The logistic differential equation models that kind of limited growth.

The logistic equation

The right side is 00 when P=0P = 0 and when P=LP = L, so those are the equilibrium solutions. For 0<P<L0 < P < L both factors are positive and PP increases. For P>LP > L the factor L−PL - P is negative and PP decreases. The right side and its derivative with respect to PP are continuous, so no solution curve crosses P=LP = L. A solution's slope approaches 00 only as PP approaches 00 or LL. So every solution with P(0)>0P(0) > 0 approaches LL:
lim⁡t→∞P(t)=L.\lim_{t \to \infty} P(t) = L.
As a function of PP, the rate kP(L−P)kP(L - P) is a downward-opening parabola with zeros at 00 and LL. Its largest value is at the vertex, P=L2P = \frac{L}{2}. So a population that starts between 00 and L2\frac{L}{2} grows fastest when it reaches half the carrying capacity. One that starts between L2\frac{L}{2} and LL grows fastest at the start.
Differentiating with the chain rule gives the concavity:
d2Pdt2=k(L−2P) dPdt=k2P(L−P)(L−2P).\begin{aligned} \frac{d^2P}{dt^2} &= k(L - 2P)\,\frac{dP}{dt} \\[4pt] &= k^2 P(L - P)(L - 2P). \end{aligned}
For 0<P<L20 < P < \frac{L}{2} this is positive, for L2<P<L\frac{L}{2} < P < L it's negative, and for P>LP > L it's positive again. A solution that starts between 00 and L2\frac{L}{2} is concave up until it reaches L2\frac{L}{2} and concave down after, so its graph has an inflection point where P=L2P = \frac{L}{2}. A solution that starts between L2\frac{L}{2} and LL is concave down throughout. One that starts above LL decreases and is concave up.

Relative growth rate

A logistic model is often written as dPdt=rP(1−PL)\frac{dP}{dt} = rP\left(1 - \frac{P}{L}\right). Here r=kLr = kL, and dividing by PP shows what it means. The relative growth rate dP/dtP=r(1−PL)\frac{dP/dt}{P} = r\left(1 - \frac{P}{L}\right) is close to rr when PP is small, and it falls to 00 as PP approaches LL.